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, coordinated changes in flower and bill shape stimulated the formation of new species of hummingbirds and plants. Due to this exceptional evolutionary pattern, as many as 140 hummingbird species can coexist in a specific re evolutionary tree shows ancestral hummingbirds splitting from insectivorous swifts (family Apodidae) and treeswifts (family Hemiprocnidae) about 42 million years ago, probably in Eurasia. One key evolutionary factor appears to be an altered taste receptor that enabled hummingbirds to seek nectar. By 22 million years ago the ancestral species of current hummingbirds became established in South America, w a particularly rich environment for hummingbird evolution because diversification occurred simultaneously with mountain uplift over the past 10 million years. Hummingbirds remain in dynamic diversification inhabiting ecological regions across South America, North America, and the Caribbean, indicating an enlarging evolutionary radiatithan males. Conversely, the Lophomithini clade displays very little size dimorphism; males and females are similar in size. Sexual dimorphisms in bill size and shape are also present between male and female hummingbirds, where in many clades, females have longer, more curved bills favored for accessing nectar from tall flowers. For males and females of the same size, ingbird and the sicklebills, are co-evolved with a small number of flower species. However, even in the most specialized hummingbird-plant mutualisms the number of food plant lineages of the individual hummingbird species increases with time. The bee hummingbird (Mellisuga helenae) – the world's smallest bird – evolved to dwarfism likely because it had to compete with long-billed hummingbirds having an advantage for nectar foraging from specialized flowers, consequently leading the bee hummingbird to more successfully compete for flowerd by hummingbirds produce flowers in shades of red, orange, and bright pink, though the birds will take nectar from flowers of other colors as well. Hummingbirds can see wavelengths into the near-ultraviolet, but hummingbird-pollinated flowers do not reflect these wavelengths as many insect-pollinated flowers do. This narrow color spectrum may render hummingbird-pollinated flowers relatively inconspicuous to most insects, thereby reducindue to constraints imposed by courtship because mating displays of male hummingbirds require complex aerial maneuvers. Males tend to be smaller than females, allowing conservation of energy to forage competitively and participate more frequently in courtship. Thus, sexual selection will favor smallel needs of visual processing while in rapid flight or hovering by development of an exceptionally dense array of retinal neurons allowing for increased spatial resolution in the lateral and frontal visual fields. Morphological studies showed that neuronal hypertrophy, relatively the largest in any bird, exists in a brain region called the pretectal nucleus lentiformis mesencephali (or nucleus of the optic tract in mammals) responsible for refining dynamic visual processing while hovering and during rapid flight. The enlargement of this brain region responsible for visual processing indicates enhanced ability for perception and processing of fast-moving visual stimuli which hummingbirds encoisual fields, responding to even minimal motion in any direction by reorienting themselves in midflight. Hummingbirds' visual sensitivity allows them to precisely hover in place while in complex and dynamic natural environments, functions enabled by the lentiform nucleus which is tuned to fast-pattern velocities, enabling highly tuned control and collisr male hud to be larger, requiring more energy, with longer beaks that allow for more effective reach into crevices of tall flowers for nectar. Thus females are better at foraging, acquiring flower nectar, and supporting the energy demands of their larger body size. Directional selection will thus favor the larger hummingbirds in terms of acquiring fmented. Though scientists theorize that hummingbirds originated in South America, where species diversity is greatest, possible ancestors of extant hummingbirds may have lived in parts of Europe to what is southerecies of hummingbirds are described, depending on taxonomic viewpoint, divided into two subfamilies, the hermits (subfamily Phaethornithinae, 34 species in six genera), and the typical hummingbirds (subfamily Trochilinae, all the others). However, recent phylogenetic analyses suggest that this division is slightly inaccurate, and that there are nine major clades of hummingbirds: the topazes and jacobins, the hermits, the mangoes, the coquettes, the brilliants, the giant hummingbird (Patagona gigas), the mountain-gems, the bees, and the emeralds. The topazes and jacobins combined have the oldest split with the rest of the hummingbirds. The hummingbird family has the second-greatest number of species of any bird family (after thewn from the Pleistocene of Brazil and the Bahamas; however, neither has yet been scientifically described, and fossils and subfossils of a few extant species are known. Until recently, older fossils had not been securely identifiable as those of d Mayr of the Senckenberg Museum in Frankfurt am Main identified two 30-million-year-old hummingbird fossils. The fossils of this primitive hummingbird species, named Eurotrochilus inexpectatus ("unexpected European hummingbird"), had been sitting in a museum drawer in Stuttgart; they had been unearthed in a clay pit at Wiesloch–Frauenweiler, south of Heidelberg, Germany, and because it was assumed that hummingbirds never occurred ose of this sexual bill dimorphism is that the selective forces from competition for nectar between the sexes of each species are what drive the sexual dimorphism. Depending on which sex holds territory in the species, it is advantageous for the other sex to have a longer bill and be able to feed on a wide variety of flowers, decreasing intraspecific competition. For example, in species of hummingbirds where males have longer bills, males do not hold a specific territory and have a lek mating system. In species where males have shorter bills than females, males defend their resources and therefore females must have a longer bill in order aphic region, hummingbird clades co-evolved with nectar-bearing plant clades, affecting mechanisms of pollination. The same is true for the sword-billed hummingbird (Ensifera ensifera), one of the morphologically most extreme species,
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